Aja, Iris, Department of Physiology, School of Medicine and Nursing, University of the Basque Country – UPV/EHU, 48940 Leioa, Spain - University of Bordeaux, ISVV, EA 4577, USC 1366 INRA, Unité de Recherche Œnologie, 210 Chemin de Leysotte, F-33882 Villenave d’Ornon, France